References

Seismic energy transmits most efficiently between 10-40 Hz - in the same range as the fundamental frequency and 2nd harmonic of an elephant rumble. It turns out that when an elephant rumbles a replica of the airborne sound is also transmitted through the ground. Elephant sounds have been measured as traveling at about 309 meters per second through air and at about 248-264 meters per second through the ground.

Experiments carried out by Caitlin O'Connell and colleagues have shown that elephants are able to pick up these seismic signals, to orient in the direction that the vibrations come from and even to respond to them appropriately.

Elephants may be able to detect these seismic vibrations, or rayleigh waves, through two possible means: bone conduction and the use of massive ossicles of their middle ears or, possibly, by mechano-receptors in the toes, feet or trunk that are sensitive to vibrations.

The tip of an elephant's trunk has layers of cells called Pacinian corpuscles that are extremely sensitive to vibrations and are thought to be able to detect movement as subtle as Brownian motion. Pacinian corpuscles have also been found in the elephant foot - concentrated in the front and back (toes and heel area) dermal layer. Movements or vibrations deform the layers of Pacinian corpuscles, sending a nerve signal to the brain. Although these corpuscles are found in other mammals, too, they are particularly densely packed in the tip of an elephant's trunk.

You can find an interesting article about seismic signals among elephants here, and numerous others through a Google search on “Seismic signals from elephants”. You will enjoy watching Dr. Caitlin E. O'Connell talk about this exciting topic on TEDx Talks in 2013, and an episode of Deep Look (KQED San Francisco, presented by PBS) called "How Elephants Listen... With Their Feet", from July 2018. Both films are embedded below.

Try using the Search Portal in the The Elephant Ethogram: A Library of African Elephant Behavior  to look for Behaviors that are likely to be involved in seismic communication. In the Combined Dropdown Search under Communication Mode, select "Seismic" to list many of the Behaviors and Constellations that involve seismic communication.